Alex, Marty, Gloria and Melman are still fighting to get home to their beloved Big Apple. Their journey takes them through Europe where they find the perfect cover: a traveling circus, which they reinvent - Madagascar style.
Spoiled by their upbringing with no idea what wild life is really like, four animals from New York Central Zoo escape, unwittingly assisted by four absconding penguins, and find themselves in Madagascar, among a bunch of merry lemurs
The Dragon Warrior has to clash against the savage Tai Lung as China's fate hangs in the balance: However, the Dragon Warrior mantle is supposedly mistaken to be bestowed upon an obese panda who is a tyro in martial arts.
In order to power the city, monsters have to scare children so that they scream. However, the children are toxic to the monsters, and after a child gets through, two monsters realize things may not be what they think.
Manny, Sid, and Diego discover that the ice age is coming to an end, and join everybody for a journey to higher ground. On the trip, they discover that Manny, in fact, is not the last of the woolly mammoths.